In the Antarctic, after an expedition with Dr. Davis McClaren, the sled dog trainer Jerry Shepherd has to leave the polar base with his colleagues due to the proximity of a heavy snow storm. He ties his dogs to be rescued after, but the mission is called-off and the dogs are left alone at their own fortune. For six months, Jerry tries to find a sponsor for a rescue mission.
When 11-year-old Riley moves to a new city, her Emotions team up to help her through the transition. Joy, Fear, Anger, Disgust and Sadness work together, but when Joy and Sadness get lost, they must journey through unfamiliar places to get back home.
Ronia lives happily in her father's castle until she comes across a new playmate, Birk, in the nearby dark forest. The two explore the wilderness, braving dangerous Witchbirds and Rump-Gnomes. But when their families find out Birk and Ronia have been playing together, they forbid them to see each other again. Indeed, their fathers are competing robber chieftains and bitter enemies. Now the two spunky children must try to tear down the barriers that have kept their families apart for so long.